In payoutStakers() I see that Era argument needs to between current_era and history_depth. So are unclaimed payouts for very old eras effectively lost? If they are unclaimable, then are those funds moved to treasury? Thanks.

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correct, when new era triggered the underlying storage like ErasStakers* and ErasValidator* will be removed and unclaimable. Staking reward is usually a model of inflation(mint/deposit) so no transfer/move required

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